Winner, Female Fifty Fathoms. Humpback whales in shallow water. Turks and Caicos Islands. #

Overall Winner. A paper nautilus drifts on a piece of ocean debris at night, surrounded by heavy sediment. Philippines. #

Second Place, Wildlife. A gentoo penguin, the fastest penguin species in the world, charges across the water in Antarctica. #

Second Place, Overall. A crab sits in the center of a sea anemone as it sways in ocean current. Philippines. #

Highly Commended, Adventure. A freediver appears to be flying in the calm blue waters of the Aegean Sea, in Greece. #

Highly Commended, Fine Art. A whitemouth moray eel’s intricate body fills the image’s entire background. Réunion Island. #

Highly Commended, Fine Art. A curious spider squat lobster checks out the photographer’s camera lens. Philippines. #

Highly Commended, Conservation (Hope). A fever of mobula rays swim peacefully in the shallow waters of the Gulf of California. #

Highly Commended, Conservation (Hope). A green sea turtle briefly poses for an angelic portrait in Maui, Hawaii, where sea turtles enjoy strong legal protections. #

Third Place, Fine Art. Hidden yet unmistakable, a northern gannet stares through a cloud of bubbles after diving in pursuit of fish. Scotland. #

Highly Commended, Adventure. A free diver is surrounded by a shiver of grey reef sharks in French Polynesia. #

Third Place, Conservation (Hope). During low tide, a coral reef is perfectly mirrored on the surface, off Mayotte Island. #

Highly Commended, Fine Art. A porcelain crab sits on top of a sea pen, its constant companion, in the Philippines. #

Highly Commended, Fine Art. A curious group of Steller sea lions circles the photographer. Canada. #

Third Place, Adventure. A scuba diver explores the underside of a gigantic iceberg in East Greenland. #

Winner, Wildlife. A lizardfish’s open mouth reveals its last meal. Philippines. #

Winner, Fine Art. A whale shark and its entourage of remoras is attracted by the bright lights of fishermen. The Maldives. #

Winner, Conservation (Impact). A polar-bear cub contends with the fragility of melting ice near Svalbard, Norway. #

Highly Commended, Human Connection. Off Alor Island, Indonesia, local children play in the water with their homemade wooden goggles held together by fishing line. #

Highly Commended, Fine Art. Rain and wind, paired with the light shining down on the spray, create a dramatic scene off Queensland, Australia. #